Maybe the situation here would be better had they had more interaction with the people at the shelter, however they were not there very long (a couple of weeks at the most).
Difficult to know really.
The kitten I adopted from the place had been there her whole life of 5 months and they were going to be rid of her the next day but ... along came I!
They get lots of petting and touches here. Its just the one with the PTSD that is difficult.
Another idea is to use a flea comb on them.
The one that bites just loves to be flea combed! He'll come and get you and give you that "look" as to say, "Comb me now!". I responded to the little dictator at once after I get that "look".
If I say the word "Comb" he comes a running! That is how much he enjoys being combed (plus the added benefit associated with it: no fleas, hairballs in check, etc.). 
Definitely worth a try and the comb you can buy for a few bucks at the most!
Best of luck!!
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