Laughing says, "I'm only play-biting you, not real-biting you."
BY HANNAH SEO MAY 18, 2021

When we think of all the creatures who laugh, humans and hyenas are probably the only ones that come to mind. But recently, scientists combed through the literature and found that at least 65 diverse species of animals produce vocalizations that could be analogous to a human chuckle. And hyenas are not one of them.
Human laughter is a vocalization that signals play, which is an important and complex social interaction, according to anthropologists and cognitive scientists in a recent paper. But humans are certainly not the only animals that play, so the researchers parsed through existing data to see which other animals produce play vocalizations, and whether those sounds were exclusive to play.
The 65 snickering species identified vary from our close ape relatives, like chimpanzees or bonobos, to more surprising mammals like slow lorises, sea lions, and orcas. But non-mammals made the list too, specifically three birds: kea parrots, parakeets, and Australian magpies. Hyenas didnt make the cut, on the other hand, because their cackles are decidedly not for play. While 28 species out of the 65 had sounds that were exclusive to play, its still unclear whether the vast majority of giggly creatures have specific sounds for play purposes. The findings were published in Bioacoustics in April.
Sixty-five might seem like a high number, but its possible that many more species produce laugh-like noises. It could be that a lot of animals do have play vocalizations [and] theyre just really quiet, paper co-author and UCLA anthropology graduate student Sasha Winkler told Ars Technica. Until scientists try to comprehensively document play sounds in all animals, we wont know which creatures are or are not capable of laughter.
More:
https://www.popsci.com/animals/animal-laughter/